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- In the late 1950s, timid sea monsterchild Luca Paguro herds goatfish off the coast of the Italian town ofPortorosso. His parents forbid him from approaching the surface, fearing that hemight be hunted by humans.
- One day, Luca meets Alberto Scorfano, a fellowseamonster child who lives alone in an abandoned tower on dry land.When Alberto encourages Luca to venture out of theocean, it is revealed that sea monsters turn into humans as long as they aredry. Luca and Alberto begin hanging out and become friends, and they wish toown a Vespa and travel around the world.Upon discovering their son'sactions, Luca's parents decide to send him to live in the deep with his uncle.
- Upon discovering their son's actions, Luca's parentsdecide to send him to live in the deep with his uncle. Inretaliation, Luca runs away with Alberto to hide out in Portorosso. The boysrun afoul of Ercole Visconti, the local bully and five-time champion of thePortorosso Cup, a children's triathlon of swimming, pasta-eating, andbiking. When Ercole tries to soak Luca in a fountain, Giulia Marcovaldo, ayoung girl, stops him. In hopes of winning the money needed for a Vespa, theboys team up with Giulia for the triathlon.Giulia invites them to stay at herhouse and introduces her fisherman father, Massimo, who holds grudges towards sea monsters.
- Giulia and Luca bond over their love of learning, making Albertojealous. When Luca shows interest in attending school, Alberto intentionally reveals his sea monster form to Giulia to prevent it. Unwilling to give himselfup, Luca feigns surprise to be with Giulia, and a betrayed Alberto flees asErcole's gang show up and try to hunt him.Shortly after, Giulia finds out that Luca is a sea monster as well, and sends him away for his safety.Luca attempts to reconcile with Alberto, who is revealed to have been abandoned by his fatherlong ago. He promises to win the Vespa to rebuild their friendship.
- The triathlon begins with Luca and Giulia competing separately.Luca manages to complete the swimming race and pasta-eating contest withoutrevealing himself, but rain begins during the bike race. Alberto shows up togive him an umbrella, only to be kicked by Ercole to the wet ground andexposed, to everyone's shock.They resume biking, with Ercole now trying toharpoon them for money, which is stopped by Giulia smashing her bike againsthis. Luca and Alberto unintentionally cross the finish line, but turn back tohelp Giulia. When Ercole and other townsfolk confront them, Massimo stands upfor them and insists that they have won. A few of the townsfolk are revealed tobe sea monsters as well and Luca is finally reunited with his family. Thehumans welcome the sea monsters, to the exception of Ercole, whose prejudiceturns him into an outcast.
- Luca and Alberto buy a Vespa, but Alberto sells it to get atrain ticket for Luca, allowing him to go to school in Genoa with Giulia.Luca's family, Massimo, and Alberto see Luca and Giulia off at the trainstation, where they all promise to stay in touch. During the credits, Lucameets Giulia's mother, whereas Massimo adopts Alberto.
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