Chris Rock makes a joke about Will Smiths wife Jada Smith that references Jada's alopecia.
Jada & Will Smith
The entire audience laughs because they find it funny, but Jada doesn't and she looks very upset.
Will Smith does something surprising
Chris Rock laughs because he sees the audience laugh at his joke.
Will Smith
Keep my wife's name out your mouth!
Will Smith sees that his wife is upset by the joke, and that she doesn't find it very funny. So he stands up and walks toward Chris Rock very angrily. 
Will Smith walks towards Chris and slaps him in the face very hard. The entire audience goes quiet, then he walks away back to his seat.
*SLAPS*
As Will Smith goes to sit down he says something to Chris Rock that left the audience very surprised, and speechless.
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